1. Aquatic Ecology Laboratory Department of Evolution, Ecology, and Organismal Biology The Ohio State University 1314 Kinnear Road 230 Research Center Columbus Ohio 43212 USA
2. U.S. Geological Survey, Wisconsin Cooperative Fishery Research Unit College of Natural Resources University of Wisconsin–Stevens Point 800 Reserve Street Stevens Point Wisconsin 54481 USA
3. U.S. Geological Survey Wisconsin Cooperative Fishery Research Unit College of Natural Resources University of Wisconsin–Stevens Point 800 Reserve Street Stevens Point Wisconsin 54481 USA
4. Department of Evolution, Ecology, and Organismal Biology The Ohio State University 318 West 12th Avenue 300 Aronoff Laboratory Columbus Ohio 43210 USA