1. Ohio Department of Natural Resources Division of Wildlife Inland Fisheries Research Unit 10517 Canal Road SE Hebron Ohio 43025 USA
2. Aquatic Ecology Laboratory Department of Evolution, Ecology, and Organismal Biology The Ohio State University 1314 Kinnear Road Columbus Ohio 43212 USA
3. Ohio Department of Natural Resources, Division of Wildlife 2045 Morse Road Columbus Ohio 43229 USA