1. Guangxi Key Laboratory of Plant Conservation and Restoration Ecology in Karst Terrain, Guangxi Institute of Botany Guangxi Zhuang Autonomous Region and Chinese Academy of Sciences Guilin 541006 China
2. The Royal Botanic Gardens Kew, Identification & Naming Department Richmond Surrey TW9 3AE United Kingdom
3. Singapore Botanic Gardens National Parks Board 1 Cluny Road Singapore 259569 Singapore
4. International Center for Tropical Botany, Institute of Environment, Department of Biological Sciences Florida International University 11200 SW 8th Street, OE 148 Miami Florida 33199 U.S.A.