1. Natural Resource Ecology Lab, Colorado State University, Fort Collins, CO 80526; USA, in Cooperation with the US Geological Survey; Fort Collins Science Center; Fort Collins Colorado USA
2. Natural Resource Ecology Lab; Department of Ecosystem Science and Sustainability; Colorado State University, Fort Collins, CO 80526, USA, in Cooperation with the US Geological Survey; Fort Collins Science Center; Fort Collins Colorado USA
3. Fort Collins Science Center; U.S. Geological Survey; Fort Collins CO 80526 USA