1. Department of Systematic Botany; Institute of Plant Sciences; University of Göttingen; Untere Karspüle 2 37073 Göttingen Germany
2. Department of Biology; Faculty of Science; Burapha University; 169 Long-Hard Bangsaen Road, Saen Sook Sub-district Mueang District, Chonburi 20131 Thailand
3. Muséum National d'Histoire Naturelle; Dept. Systématique et Evolution; C.P. 39, 57 rue Cuvier 75231 Paris cedex 05 France
4. Netherlands Centre for Biodiversity Naturalis; section NHN; Leiden University; P.O. Box 9514 2300 RA Leiden The Netherlands