1. Department of Wildlife, Fish, and Environmental Studies Swedish University of Agricultural Sciences Umeå SE‐90183 Sweden
2. Norwegian Institute for Nature Research Sognsveien 68 Oslo NO‐0855 Norway
3. Department of Zoology Centre for African Conservation Ecology Nelson Mandela University Port Elizabeth 6031 South Africa
4. Copernicus Institute of Sustainable Development Environmental Sciences Utrecht University Utrecht 3548 The Netherlands
5. Norwegian Institute for Nature Research PO Box 5685 Torgard Trondheim NO‐7485 Norway
6. Department of Ecology Swedish University of Agricultural Sciences Grimsö Wildlife Research Station RiddarhyttanSE‐73993 Sweden
7. Department of Forestry and Wildlife Management Inland Norway University of Applied Sciences Koppang NO‐2480 Norway