1. Netherlands Centre for Biodiversity Naturalis (section NHN), Leiden University,; P.O. Box 9514, RA Leiden, 2300 The Netherlands,
2. Hortus botanicus Leiden,; P.O. Box 9516, RA Leiden, 2300 The Netherlands;,
3. Laboratory of Plant Systematics, Institute of Botany and Microbiology, K.U. Leuven,; Kasteelpark Arenberg 31, Leuven, 3001 Belgium