1. Key Laboratory of Agro-ecological Processes in Subtropical Region, Institute of Subtropical Agriculture; Chinese Academy of Sciences; Changsha Hunan 410125 China
2. Huanjiang Observation and Research Station for Karst Ecosystems; Chinese Academy of Sciences; Huanjiang Guangxi 547100 China
3. College of Environmental Science and Engineering; Guilin University of Technology; Guilin 541004 China
4. College of Resources and Environment; Hunan Agricultural University; Changsha Hunan 410128 China
5. Graduate University of Chinese Academy of Sciences; Beijing 100049 China