1. Aix Marseille Univ.; Univ Avignon, CNRS, IRD, UMR 7263; Mediterranean Institute of Marine and Continental Biodiversity and Ecology (IMBE); Station Marine d'Endoume Marseille France
2. Aix Marseille Univ.; CNRS, UMR 7288; IBDM; Marseille France
3. Department of Biological Sciences; University of Alberta; Edmonton, Alberta T6G 2E9 Canada
4. Department of Earth and Environmental Sciences, Paleontology and Geobiology; Ludwig-Maximilians-Universität München; Richard-Wagner Straße 10 80333 Munich Germany
5. GeoBio-Center; Ludwig-Maximilians-Universität München; Munich Germany
6. Bavarian State Collection for Paleontology and Geology; Munich Germany