1. Smithsonian Conservation Biology Institute, National Zoological Park1500 Remount Rd.Front RoyalVA 22630USA
2. Faculty of Technology, Natural Sciences, and Maritime Sciences, Department of Natural Sciences and Environmental Health, University College of Southeast Norway3800 Bø i TelemarkNorway
3. Wildlife Conservation Research Unit, Zoology Department, The Recanati‐Kaplan Centre, University of Oxford, Tubney HouseAbingdon RoadTubneyAbingdon, OX13 5QLUK