1. Digestive System Research Unit; Vall d'Hebron Research Institute; Passeig Vall d'Hebron 119-129 Barcelona Spain
2. Center for Integrative Genomics; University of Lausanne; Lausanne Switzerland
3. INSERM UMR 1037, CRCT, Université de Toulouse, UPS; Toulouse France
4. Université de Toulouse, UPS; Toulouse France
5. INSERM U1220; IRSD, Université de Toulouse, INSERM, INRA, ENVT, UPS; Toulouse France