1. College of Science and Engineering James Cook University Townsville Queensland 4811 Australia
2. ARC Centre of Excellence for Coral Reef Studies James Cook University Townsville Queensland 4811 Australia
3. Centre for Biological Diversity Scottish Oceans Institute University of St. Andrews St. Andrews KY16 9TH UK
4. Hawai'i Institute of Marine Biology University of Hawai'i at Mānoa 46‐007 Lilipuna Rd Kaneohe Hawai'i 96744 USA
5. Department of Biological Sciences Macquarie University Sydney New South Wales 2109 Australia