1. Aquatic Systems Research Group, Department of Geography and Environmental Sciences University of Venda Thohoyandou South Africa
2. School of Biological Sciences Queen's University Belfast Belfast UK
3. South African Institute for Aquatic Biodiversity Makhanda South Africa
4. Aquatic Systems Research Group, School of Biology and Environmental Sciences University of Mpumalanga Nelspruit South Africa
5. Wissenschaftskolleg zu Berlin ‐ Institute for Advanced Study Berlin Germany