1. Department of Crop Science College of Agricultural Sciences São Paulo State University Botucatu São Paulo 18.610‐307 Brazil
2. Postgraduate Program in Agroecology Maranhão State University São Luis Maranhão 65000‐000 Brazil
3. Federal Institute of Education, Science and Technology of Piauí Uruçuí Piauí 64860‐000 Brazil
4. Department of Biology Federal University of Maranhão São Luís Maranhão 65080‐805 Brazil
5. Institute of Biological and Environmental Sciences University of Aberdeen Aberdeen AB24 3UU UK
6. School of Biosciences University of Nottingham Sutton Bonington Campus Loughborough LE125RD UK