1. Research Centre for Ecological changes, Organismal and Evolutionary Biology Research Program; Faculty of Environmental and Biological Sciences; University of Helsinki; Helsinki Finland
2. Laboratory of Microbial Ecology; University of Lyon, University Claude Bernard Lyon 1; UMR CNRS 5557, UMR INRA 1418 VetAgro Sup Villeurbanne France
3. Finnish Museum of Natural History; Zoology Unit; University of Helsinki; Helsinki Finland