1. College of Fisheries Huazhong Agricultural University Wuhan Hubei 430070 China
2. Engineering Research Center of Green Development for Conventional Aquatic Biological Industry in the Yangtze River Economic Belt Ministry of Education Wuhan Hubei 430070 China
3. Hubei Provincial Engineering Laboratory for Pond Aquaculture Wuhan Hubei 430070 China
4. Hubei Hongshan Laboratory Wuhan Hubei 430070 China