1. Laboratory of Soft & Wet Matter; Graduate School of Life Science; Hokkaido University; Kita-10-Nishi-8 Kita-ku, Sapporo 060-0810 Japan
2. Laboratory of Soft & Wet Matter; Faculty of Advanced Life Science; Hokkaido University; Kita-10-Nishi-8 Kita-ku, Sapporo 060-0810 Japan
3. Laboratory of Soft & Wet Matter; Graduate School of Science; Hokkaido University; Kita-10-Nishi-8 Kita-ku, Sapporo 060-0810 Japan
4. Laboratory of Proteoglycan Signaling and Therapeutics; Faculty of Advanced Life Science; Hokkaido University; Kita-21-Nishi-11 Kita-ku, Sapporo 001-0021 Japan
5. Department of Sports Medicine and Joint Surgery; Graduate School of Medicine; Hokkaido University; Kita-15-Nishi-7 Kita-ku, Sapporo 060-0821 Japan
6. Biomechanics & Calorimetry Center Basel; Faculty of Medicine; University of Basel; c/o Biozentrum, Klingelbergstrasse 50-70 4056 Basel Switzerland