1. Marine Biology Institute, College of Science, Shantou University Shantou Guangdong Province China
2. Guangxi Major Laboratory of Beibu Gulf Marine Biodiversity ConservationQinzhou University Guanxi China
3. Department of Marine Science, College of OceanQinzhou University Guangxi China
4. Guanxi Academy of Science Nanning Guanxi China
5. Laboratory of Marine Biology and EcologyThird Institute of Oceanography, State Oceanic Administration Xiamen China
6. Clymene Enterprises Lakeside CA USA
7. College of Resource and EnvironmentQinzhou University Guangxi China