1. Key laboratory of Coastal Biology and Bioresource Utilization Yantai Institute of Costal Zone Research Chinese Academy of Sciences Yantai China
2. Laboratory for Marine Biology and Biotechnology Qingdao National Laboratory for Marine Science and Technology Qingdao China
3. University of Chinese Academy of Sciences Beijing China
4. Marine Science Research Institute of Shandong Province National Oceanographic Center of Qingdao Qingdao China