1. School of Biosciences, College of Life and Environmental Sciences; University of Exeter; Exeter; EX4 4PS; UK
2. National Institute for Environmental Studies; 16-2 Onogawa; Tsukuba; Ibaraki; 305-8506; Japan
3. Japan NUS Co. Ltd, 7-5-25 Nishi-Shinjyuku; Shinjyuku-ku; Tokyo; 160-0023; Japan