1. Haub School of Environment and Natural Resources University of Wyoming 804 E. Fremont Street Laramie Wyoming 82072 USA
2. Wyoming Cooperative Fish and Wildlife Research Unit Department of Zoology and Physiology University of Wyoming 1000 E. University Avenue Laramie Wyoming 82071 USA
3. Wyoming Game and Fish Department P.O. Box 1022 Thayne Wyoming 83127 USA