1. Groningen Institute for Evolutionary Life Sciences University of Groningen Nijenborg 7 9747AG Groningen The Netherlands
2. Tanzania National Parks Arusha Tanzania
3. Centre for African Ecology School of Animal, Plant and Environmental Sciences University of the Witwatersrand Wits 2050 South Africa
4. Department of Ecology and Evolutionary Biology Princeton University Princeton New Jersey 08544‐2016 USA
5. Institute of Environmental Sciences Leiden University Einsteinweg 2 2333CC Leiden The Netherlands