1. Spatial Epidemiology and Ecology Research Laboratory, Department of Geography, Emerging Pathogens Institute University of Florida Gainesville FL 32611 USA
2. Montana Fish, Wildlife, and Parks Bozeman MT 59718 USA
3. Turner Enterprises 1123 Research Drive Bozeman MT 59718 USA
4. Quantitative Disease Ecology and Conservation Laboratory, Department of Geography, Emerging Pathogens Institute University of Florida Gainesville FL 32611 USA