1. Key Laboratory of Vertebrate Evolution and Human Origins, Institute of Vertebrate Paleontology and Paleoanthropology Chinese Academy of Sciences Beijing China
2. College of Earth and Planetary Sciences University of Chinese Academy of Sciences Beijing China
3. CAS Center for Excellence in Life and Paleoenvironment Chinese Academy of Sciences Beijing China
4. Institute for Ecological Research and Pollution Control of Plateau Lakes, School of Ecology and Environmental Science Yunnan University Kunming China