1. Zoology Department, College of Science King Saud University Riyadh Kingdom of Saudi Arabia
2. Biology Department, College of Science Al Imam Mohammad Ibn Saud Islamic University (IMSIU) Riyadh Saudi Arabia
3. Groupe de Recherche en Écologie Buccale Université Laval Québec Québec Canada
4. Department of Biochemistry, College of Science King Saud University Kingdom of Saudi Arabia Riyadh