1. Odum School of Ecology; University of Georgia; Athens Georgia 30602 USA
2. Oceanography Section; Science Research Center; Kochi University; 200 Otsu, Monobe Nankoku Kochi 783-8502 Japan
3. Laboratory of Aquatic Environmental Science; Faculty of Agriculture and Marine Science; Kochi University; 200 Otsu, Monobe Nankoku Kochi 783-8502 Japan