1. State Key Laboratory of Freshwater Ecology and Biotechnology; Institute of Hydrobiology; Chinese Academy of Sciences; Wuhan 430072 China
2. University of Chinese Academy of Sciences; Beijing 100049 China
3. Key Laboratory of Aquatic Biodiversity and Conservation; Institute of Hydrobiology; Chinese Academy of Sciences; Wuhan 430072 China
4. Idaho Department of Fish and Game; Eagle Fish Health Laboratory; 1800 Trout Road Eagle Idaho 83616 USA