1. Hubbs-SeaWorld Research Institute; 2595 Ingraham Street San Diego California 92109 USA
2. National Marine Fisheries Service; Northwest Fisheries Science Center; Environmental and Fisheries Sciences Division; 2725 Montlake Boulevard East Seattle Washington 98112 USA
3. Department of Animal and Veterinary Science; University of Idaho; 875 Perimeter Drive Moscow Idaho 83844 USA