1. College of Plant Protection; Shandong Agricultural University; Tai'an Shandong 271018 China
2. Cotton Research Center; Shandong Academy of Agricultural Sciences; Jinan 250100 China
3. Institute for Genomics and Systems Biology; The University of Chicago; Chicago Illinois 60615
4. Tropical crops genetic resources institute; Danzhou Hainan 571737 China
5. Institute of Zoology; Chinese Academy of Sciences; Beijing 100101 China