1. Department of Research and ConservationParco Natura Viva ‐ Garda Zoological ParkBussolengo Italy
2. Dipartimento di Scienze Mediche VeterinarieUniversità di BolognaBologna Italy
3. Department of Biology, Chemistry and Forensic ScienceUniversity of WolverhamptonWolverhampton UK
4. Department of Anthropology and Behaviour, Ecology and Evolution Research (BEER) CentreDurham UniversityDurham UK