1. Department of Aquaculture; National Penghu University of Science and Technology; Makung City Penghu 880 Taiwan
2. Institute of Biomedical Engineering, College of Medicine and College of Engineering; National Taiwan University; Taipei 100 Taiwan
3. Penghu Marine Biology Research Center, Fisheries Research Institute, Council of Agriculture; Penghu 880 Taiwan
4. Department of Life Sciences; National Chung Hsing University; Taichung 402 Taiwan