1. Key Laboratory for Plant Diversity and Biogeography of East Asia Kunming Institute of Botany, Chinese Academy of Sciences Kunming Yunnan 650201 China
2. University of the Chinese Academy of Sciences Beijing 100049 China
3. School of Life Sciences Yunnan University Kunming Yunnan 650091 China
4. College of Pharmaceutical Science Yunnan University of Chinese Medicine Kunming Yunnan 650500 China
5. School of Integrative Plant Science, Section of Plant Biology and the L.H. Bailey Hortorium Cornell University Ithaca New York 14850 U.S.A.
6. Key Laboratory of Plant Germplasm Enhancement and Specialty Agriculture Wuhan Botanical Garden, Chinese Academy of Sciences Wuhan Hubei 430074 China
7. Central Herbarium of Uzbekistan Institute of Botany, Academy Sciences of Uzbekistan Tashkent 100125 Uzbekistan