1. Ecology Department of Biology University of Konstanz Konstanz78464Germany
2. Key Laboratory of Wetland Ecology and Environment Northeast Institute of Geography and Agroecology Chinese Academy Sciences Changchun130102China
3. Zhejiang Provincial Key Laboratory of Plant Evolutionary Ecology and Conservation Taizhou University Taizhou318000China
4. IRDIPME 911 Avenue Agropolis, BP 64501 Montpellier34394France