1. State Key Laboratory of Silkworm Genome Biology College of Sericulture Textile and Biomass Sciences Southwest University Chongqing 400715 China
2. West China Hospital Sichuan University Chengdu 610041 China
3. Botnar Research Centre, Nuffield Department of Orthopedics, Rheumatology and Musculoskeletal Sciences University of Oxford Headington Oxford OX3 7LD UK