1. Centre for Ecology & Conservation; University of Exeter; Penryn Cornwall TR 10 9EZ UK
2. Laboratory of Entomology; Wageningen University; P.O. Box 8031 Wageningen NL-6700 EH The Netherlands
3. CIRAD; UMR PVBMT; F-97410 Saint-Pierre La Réunion France
4. Community Ecology; Institute of Ecology and Evolution; University of Bern; Baltzerstrasse 6 Bern 3012 Switzerland