1. Shandong Provincial Key Laboratory of Synthetic Biology Qingdao Institute of Bioenergy and Bioprocess Technology Chinese Academy of Sciences Qingdao Shandong 266101 China
2. Shandong Energy Institute Qingdao Shandong 266101 China
3. Qingdao New Energy Shandong Laboratory Qingdao Shandong 266101 China
4. University of Chinese Academy of Sciences Beijing 100049 China
5. Marine Biology and Biotechnology Laboratory Qingdao National Laboratory for Marine Science and Technology Qingdao Shandong 266101 China