1. Key Laboratory of Experimental Marine Biology Institute of Oceanology Chinese Academy of Sciences Qingdao 266071 China
2. University of Chinese Academy of Sciences Beijing 101408 China
3. State Key Laboratory of Microbial Technology, and Marine Biotechnology Research Center Shandong University 72 Binhai Road Qingdao 266237 Shandong P. R. China
4. Laboratory for Marine Biology and Biotechnology Pilot National Laboratory for Marine Science and Technology 168 Weihai Road Qingdao 266237 Shandong P. R. China