1. Department of Biochemistry University of Oxford South Parks Road Oxford OX1 3QU UK
2. MRC Human Immunology Unit MRC Weatherall Institute of Molecular Medicine Radcliffe Department of Medicine University of Oxford Oxford OX3 9DS UK
3. Current address: InProTher Aps Ole Maaløes Vej 3 2200 København Denmark
4. Jenner Institute University of Oxford Oxford OX3 7DQ UK
5. Wellcome Trust Centre for Human Genetics University of Oxford Oxford OX3 7BN UK