1. Key Laboratory of South China Sea Fishery Resources Exploitation & Utilisation, Ministry of Agriculture of China, South China Sea Fisheries Research Institute, Chinese Academy of Fishery SciencesGuangdong Provincial Key Laboratory of Fishery Ecology and Environment Guangzhou 510300 China
2. National Engineering Research Centre for Marine AquacultureZhejiang Ocean University Zhejiang 316022 China
3. College of Marine SciencesBeibu Gulf University Qinzhou 535011 China
4. Guangxi Beibu Gulf Marine Research CenterGuangxi Academy of Sciences Nanning Guangxi 530000 China