1. CAS Key Laboratory of Mountain Ecological Restoration and Bioresource Utilization & Ecological Restoration and Biodiversity Conservation Key Laboratory of Sichuan Province Chengdu Institute of BiologyChinese Academy of Sciences Chengdu China
2. University of Chinese Academy of Sciences Beijing China
3. Ministry of Education Key Laboratory for Ecology of Tropical Islands, College of Life Sciences Hainan Normal University Haikou China