1. Shandong Key Laboratory of Radiation Oncology Shandong Cancer Hospital and Institute Shandong First Medical University Jinan Shandong P. R. China
2. Medical Science and Technology Innovation Center Shandong First Medical University Jinan Shandong P. R. China
3. Gastroenterology Research Institute and Clinical Center Shandong First Medical University Jinan Shandong P. R. China
4. Max Planck Unit for the Science of Pathogens Berlin Germany