1. Key Laboratory of Biodiversity Sciences and Ecological Engineering, Ministry of Education, College of Life Sciences; Beijing Normal University; Beijing China
2. State Key Laboratory of Vegetation and Environmental Change, Institute of Botany; Chinese Academy of Sciences; Beijing China
3. Section for Ecoinformatics and Biodiversity, Department of Bioscience; Aarhus University; Aarhus C Denmark
4. State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany; Chinese Academy of Sciences; Beijing China
5. Institute of Loess Plateau; Shanxi University; Taiyuan China
6. Center for Biodiversity Dynamics in a Changing World (BIOCHANGE); Aarhus University; Aarhus C Denmark