1. Department of Biochemistry; University of Missouri; 117 Schweitzer Hall, Columbia Missouri 65211 USA
2. Department of Veterinary Pathology; University of Missouri; Columbia Missouri USA
3. Biomedical Science and Physical Therapy Program; University of Missouri; Columbia Missouri USA
4. Department of Nutrition and Exercise Physiology; University of Missouri; Columbia Missouri USA
5. Acceleron Pharma, Inc.; Cambridge Massachusetts USA