1. Department of Gastroenterology and Metabolism; Applied Life Sciences, Institute of Biomedical and Health Sciences, Hiroshima University; Hiroshima Japan
2. Liver Research Project Center, Hiroshima University; Hiroshima Japan
3. Natural Science Center for Basic Research and Development, Hiroshima University; Hiroshima Japan
4. Laboratory for Digestive Diseases, RIKEN Center for Integrative Medical Sciences; Hiroshima Japan
5. Department of Gastroenterology and Metabolism, Graduate School of Biomedical and Health Sciences; Hiroshima University