1. Department of Plant Biology University of Illinois at Urbana‐Champaign Urbana IL 61801 USA
2. College of Earth and Environmental Sciences, MOE Key Laboratory of Western China's Environmental Systems Lanzhou University Lanzhou 730000 China
3. Appalachian Laboratory University of Maryland Center for Environmental Science Frostburg MD 21532 USA
4. School of Integrative Biology University of Illinois at Urbana‐Champaign Urbana IL 61801 USA
5. Department of Geology University of Illinois at Urbana‐Champaign Urbana IL 61801 USA
6. Program in Ecology, Evolution, and Conservation Biology University of Illinois at Urbana‐Champaign Urbana IL 61801 USA