1. Forest Sciences Center of Catalonia (CTFC) Carretera de Sant Llorenç, km.2 Solsona Catalonia E‐25280 Spain
2. Center for Ecological Research and Forestry Applications (CREAF) Cerdanyola del Vallès Catalonia E‐08193 Spain
3. Department of Agriculture and Forest Engineering (EAGROF) University of Lleida Lleida E‐25198 Spain
4. Département de Sciences Biologiques Université de Montréal CP 6128, Succursale Centre‐ville Montreal Quebec H3C 3J7 Canada
5. Independent Researcher 8 Roblyn Place Lincoln 7608 New Zealand
6. Manaaki Whenua, Landcare Research P.O. Box 40 Lincoln 7640 New Zealand
7. Department of Ecology and Evolutionary Biology University of Toronto 25 Willcocks Street Toronto Ontario M5S 3B2 Canada
8. Smithsonian Tropical Research Institute Box 0843‐03092 Balboa Ancon Panama
9. Ecology and Evolutionary Biology University of California Los Angeles California 90095 USA