1. Key Laboratory of Zoological Systematics and Evolution Institute of Zoology Chinese Academy of Sciences Beijing 100101 China
2. School of Statistics and Mathematics Yunnan University of Finance and Economics Kunming 650221 China
3. Department of Computer Science and Technology College of Information Science and Technology Beijing University of Chemical Technology Beijing 100029 China
4. College of Life Sciences Capital Normal University Beijing 100048 China
5. School of Biological Sciences University of Sydney Sydney New South Wales 2006 Australia
6. College of Life Sciences University of Chinese Academy of Sciences Beijing 100049 China