1. CAS Key Laboratory of Tropical Marine Bio-Resources and Ecology/Guangdong Key Laboratory of Marine Materia Medica/Innovation Academy of South China Sea Ecology and Environmental Engineering South China Sea Institute of Oceanology Chinese Academy of Sciences Guangzhou 510301 P. R. China
2. University of Chinese Academy of Sciences 19 Yuquan Road Beijing 100049 P. R. China
3. Southern Marine Science and Engineering Guangdong Laboratory (Guangzhou) Guangzhou 511458 P. R. China