1. CAS Key Laboratory of Tropical Marine Bio-Resources and Ecology/Guangdong Key Laboratory of Marine Materia Medica/Innovation Academy of South China Sea Ecology and Environmental Engineering South China Sea Institute of Oceanology Chinese Academy of Sciences Guangzhou 510301 P. R. China
2. Laboratory of Tropical Biological Resources of the Ministry of Education of China Hainan University Haikou 570228 P. R. China
3. Sanya Institute of Ocean Eco-Environmental Engineering Sanya 572000 P. R. China