1. CAS Key Laboratory of Tropical Marine Bio‐Resources and Ecology South China Sea Institute of Oceanology Chinese Academy of Sciences Guangzhou 510301China
2. Southern Marine Science and Engineering Guangdong Laboratory (Guangzhou) Guangzhou 511458China
3. University of Chinese Academy of Sciences Beijing 100049China
4. Institution of South China Sea Ecology and Environmental Engineering Chinese Academy of Sciences Guangzhou 510275China